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the formation of vegetable mould-第31部分

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triturating particles of rock; that there is good evidence that on 
each acre of land; which is sufficiently damp and not too sandy; 
gravelly or rocky for worms to inhabit; a weight of more than ten 
tons of earth annually passes through their bodies and is brought 
to the surface。  The result for a country of the size of Great 
Britain; within a period not very long in a geological sense; such 
as a million years; cannot be insignificant; for the ten tons of 
earth has to be multiplied first by the above number of years; and 
then by the number of acres fully stocked with worms; and in 
England; together with Scotland; the land which is cultivated and 
is well fitted for these animals; has been estimated at above 32 
million acres。  The product is 320 million million tons of earth。



CHAPTER VITHE DENUDATION OF THE LANDcontinued。



Denudation aided by recently ejected castings flowing down inclined 
grass…covered surfacesThe amount of earth which annually flows 
downwardsThe effect of tropical rain on worm castingsThe finest 
particles of earth washed completely away from castingsThe 
disintegration of dried castings into pellets; and their rolling 
down inclined surfacesThe formation of little ledges on hill…
sides; in part due to the accumulation of disintegrated castings
Castings blown to leeward over level landAn attempt to estimate 
the amount thus blownThe degradation of ancient encampments and 
tumuliThe preservation of the crowns and furrows on land 
anciently ploughedThe formation and amount of mould over the 
Chalk formation。

We are now prepared to consider the more direct part which worms 
take in the denudation of the land。  When reflecting on sub…aerial 
denudation; it formerly appeared to me; as it has to others; that a 
nearly level or very gently inclined surface; covered with turf; 
could suffer no loss during even a long lapse of time。  It may; 
however; be urged that at long intervals; debacles of rain or 
water…spouts would remove all the mould from a very gentle slope; 
but when examining the steep; turf…covered slopes in Glen Roy; I 
was struck with the fact how rarely any such event could have 
happened since the Glacial period; as was plain from the well…
preserved state of the three successive 〃roads〃 or lake…margins。  
But the difficulty in believing that earth in any appreciable 
quantity can be removed from a gently inclined surface; covered 
with vegetation and matted with roots; is removed through the 
agency of worms。  For the many castings which are thrown up during 
rain; and those thrown up some little time before heavy rain; flow 
for a short distance down an inclined surface。  Moreover much of 
the finest levigated earth is washed completely away from the 
castings。  During dry weather castings often disintegrate into 
small rounded pellets; and these from their weight often roll down 
any slope。  This is more especially apt to occur when they are 
started by the wind; and probably when started by the touch of an 
animal; however small。  We shall also see that a strong wind blows 
all the castings; even on a level field; to leeward; whilst they 
are soft; and in like manner the pellets when they are dry。  If the 
wind blows in nearly the direction of an inclined surface; the 
flowing down of the castings is much aided。

The observations on which these several statements are founded must 
now be given in some detail。  Castings when first ejected are 
viscid and soft; during rain; at which time worms apparently prefer 
to eject them; they are still softer; so that I have sometimes 
thought that worms must swallow much water at such times。  However 
this may be; rain; even when not very heavy; if long continued; 
renders recently…ejected castings semi…fluid; and on level ground 
they then spread out into thin; circular; flat discs; exactly as 
would so much honey or very soft mortar; with all traces of their 
vermiform structure lost。  This latter fact was sometimes made 
evident; when a worm had subsequently bored through a flat circular 
disc of this kind; and heaped up a fresh vermiform mass in the 
centre。  These flat subsided discs have been repeatedly seen by me 
after heavy rain; in many places on land of all kinds。

On the flowing of wet castings; and the rolling of dry 
disintegrated castings down inclined surfaces。When castings are 
ejected on an inclined surface during or shortly before heavy rain; 
they cannot fail to flow a little down the slope。  Thus; on some 
steep slopes in Knole Park; which were covered with coarse grass 
and had apparently existed in this state from time immemorial; I 
found (Oct。 22; 1872) after several wet days that almost all the 
many castings were considerably elongated in the line of the slope; 
and that they now consisted of smooth; only slightly conical 
masses。  Whenever the mouths of the burrows could be found from 
which the earth had been ejected; there was more earth below than 
above them。  After some heavy storms of rain (Jan。 25; 1872) two 
rather steeply inclined fields near Down; which had formerly been 
ploughed and were now rather sparsely clothed with poor grass; were 
visited; and many castings extended down the slopes for a length of 
5 inches; which was twice or thrice the usual diameter of the 
castings thrown up on the level parts of these same fields。  On 
some fine grassy slopes in Holwood Park; inclined at angles between 
8 degrees and 11 degrees 30 seconds with the horizon; where the 
surface apparently had never been disturbed by the hand of man; 
castings abounded in extraordinary numbers:  and a space 16 inches 
in length transversely to the slope and 6 inches in the line of the 
slope; was completely coated; between the blades of grass; with a 
uniform sheet of confluent and subsided castings。  Here also in 
many places the castings had flowed down the slope; and now formed 
smooth narrow patches of earth; 6; 7; and 7。5 inches in length。  
Some of these consisted of two castings; one above the other; which 
had become so completely confluent that they could hardly be 
distinguished。  On my lawn; clothed with very fine grass; most of 
the castings are black; but some are yellowish from earth having 
been brought up from a greater depth than usual; and the flowing…
down of these yellow castings after heavy rain; could be clearly 
seen where the slope was 5 degrees; and where it was less than 1 
degree some evidence of their flowing down could still be detected。  
On another occasion; after rain which was never heavy; but which 
lasted for 18 hours; all the castings on this same gently inclined 
lawn had lost their vermiform structure; and they had flowed; so 
that fully two…thirds of the ejected earth lay below the mouths of 
the burrows。

These observations led me to make others with more care。  Eight 
castings were found on my lawn; where the grass…blades are fine and 
close together; and three others on a field with coarse grass。  The 
inclination of the surface at the eleven places where these 
castings were collected varied between 4 degrees 30 seconds and 17 
degrees 30 seconds; the mean of the eleven inclinations being 9 
degrees 26 seconds。  The length of the castings in the direction of 
the slope was first measured with as much accuracy as their 
irregularities would permit。  It was found possible to make these 
measurements within about of an inch; but one of the castings was 
too irregular to admit of measurement。  The average length in the 
direction of the slope of the remaining ten castings was 2。03 
inches。  The castings were then divided with a knife into two parts 
along a horizontal line passing through the mouth of the burrow; 
which was discovered by slicing off the turf; and all the ejected 
earth was separately collected; namely; the part above the hole and 
the part below。  Afterwards these two parts were weighed。  In every 
case there was much more earth below than above; the mean weight of 
that above being 103 grains; and of that below 205 grains; so that 
the latter was very nearly double the former。  As on level ground 
castings are commonly thrown up almost equally round the mouths of 
the burrows; this difference in weight indicates the amount of 
ejected earth which had flowed down the slope。  But very many more 
observations would be requisite to arrive at any general result; 
for the nature of the vegetation and other accidental 
circumstances; such as the heaviness of the rain; the direction and 
force of the wind; &c。; appear to be more important in determining 
the quantity of the earth which flows down a slope than its angle。  
Thus with four castings on my lawn (included in the above eleven) 
where the mean slope was 7 degrees 19 seconds; the difference in 
the amount of earth above and below the burrows was greater than 
with three other castings on the same lawn where the mean slope was 
12 degrees 5 seconds。

We may; however; take the above eleven cases; which are accurate as 
far as they go; and calculate the weight of the ejected earth which 
annually flows down a slope having a mean inclination of 9 degrees 
26 seconds。  This was done by my 
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